Ecology Solution

A + + L + T + + + + S + + + N 
R + + + A + R T + U + + + E + 
D + + + + D R O O + + + R + + 
N + + + + E I U P + + I + + + 
U + + + S + D T + I T + + S + 
T R + E + I + + R I C + + A + 
+ E D + C + + + C E + A + V + 
+ T + E + + + Y + + T + L A + 
+ A D T A I G A R + + N + N + 
+ W + + + + + + + A + + I N + 
+ H + + + + + + + + U + + A + 
+ S G R A S S L A N D T + + + 
+ E C I N A E C O + + + S + + 
+ R + + + + + + + + + + + E + 
+ F + + + + + + + + + + + + + 

(Over,Down,Direction) 
DECIDUOUS(3,9,NE)
DESERT(3,7,NE)
ESTUARY(14,14,NW)
FRESHWATER(2,15,N)
GRASSLAND(3,12,E)
INTERTIDAL(13,10,NW)
NERITIC(15,1,SW)
OCEANIC(9,13,W)
SAVANNA(14,5,S)
TAIGA(4,9,E)
TROPICAL(6,1,SE)
TUNDRA(1,6,N)